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This low concentration of acetate is attributed to the intermolecular neutralization of the N-terminal amino group and the -amino group of lysine by carboxylate groups on the C-terminal or on the side chain of the aspartate and glutamate residues

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US patent 2413493, Bitler WP, Flisik AC, Leonard N, "Synthesis of isomer-free benzyl methyl acetoacetic methyl ester", published 31 December 1946, assigned to Kay Fries Chemicals Inc